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Selection Guide

Cre Line Selection Guide

Choosing the right Cre driver line is critical for successful conditional knockout experiments. The wrong Cre can lead to unexpected phenotypes, embryonic lethality, or failure to delete in your tissue of interest.

This guide covers key selection criteria, major Cre line categories, and resources to help you identify the optimal driver for your conditional knockout project.

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Key Selection Criteria

Tissue Specificity

Does the Cre driver express exclusively in your tissue of interest? Consider both the promoter used and actual expression data from published characterization studies.

  • Review published expression patterns and recombination maps
  • Consider both intended and unintended expression sites
  • Account for embryonic vs adult expression differences
  • Verify specificity in your strain background

Expression Timing

When does Cre expression begin? Early embryonic expression may cause deletion before your tissue of interest differentiates, while late expression may miss critical developmental windows.

  • Match Cre expression timing to your experimental needs
  • Consider developmental stage of target tissue
  • Use inducible Cre for precise temporal control
  • Account for Cre expression onset vs full activity

Recombination Efficiency

How completely does the Cre driver recombine floxed alleles in target tissues? Variable efficiency can create mosaic deletion patterns that complicate phenotype interpretation.

  • Review published recombination efficiency data
  • Consider using reporter alleles to assess efficiency
  • Higher efficiency is generally preferred
  • Account for variation between individual animals

Background Recombination

Does the Cre show any activity outside the intended tissue? Even low level leaky expression can confound results if your gene is essential in other tissues.

  • Check for germline recombination (especially with female Cre carriers)
  • Review reports of unexpected tissue recombination
  • Consider Cre toxicity in expressing tissues
  • Assess mosaicism in target and non target tissues

Major Cre Line Categories

Ubiquitous Cre Lines

Express Cre in all tissues, useful for generating global knockouts from conditional alleles.

Examples: CMV Cre (early embryonic, all tissues), EIIa Cre (early embryonic), Actin Cre (broadly expressed), Sox2 Cre (epiblast, germline)

Tissue Specific Cre Lines

Express Cre in defined tissue types based on endogenous promoter specificity.

Examples: Albumin Cre (hepatocytes), Nestin Cre (neural progenitors), LysM Cre (myeloid cells), CD4 Cre (T lymphocytes), Villin Cre (intestinal epithelium)

Cell Type Specific Cre Lines

Express Cre in specific cell populations within tissues.

Examples: CamKII Cre (forebrain neurons), Adipoq Cre (mature adipocytes), Myh6 Cre (cardiomyocytes), Pdx1 Cre (pancreatic cells), Col2a1 Cre (chondrocytes)

Developmental Stage Cre Lines

Express Cre at specific developmental timepoints.

Examples: Shh Cre (sonic hedgehog expressing cells), Wnt1 Cre (neural crest), Pax3 Cre (dermomyotome), Tie2 Cre (endothelial/hematopoietic)

Inducible Cre Systems

Inducible Cre lines provide temporal control over gene deletion, enabling deletion in adult animals or at specific developmental timepoints.

CreERT2 (Tamoxifen Inducible)

Cre fused to modified estrogen receptor. Inactive until tamoxifen administration. Most widely used inducible system.

Advantages

  • Tight temporal control
  • Wide variety of tissue specific versions available
  • Reversible (deletion is permanent, but Cre activity requires continued tamoxifen)

Considerations

  • Tamoxifen has estrogenic effects
  • Efficiency varies with dose and timing
  • Some leakiness possible without induction

Tet Inducible Cre

Cre expression controlled by doxycycline. Can be either Tet On (dox activates) or Tet Off (dox represses).

Advantages

  • Reversible control of Cre expression
  • Dose dependent expression levels
  • No estrogen related effects

Considerations

  • Requires two transgenes (tTA/rtTA and TetO Cre)
  • Background activity varies by line
  • Doxycycline has some biological effects

Cre Line Selection Workflow

1

Step 1: Define Your Requirements

Specify the tissue(s), cell type(s), and timing for gene deletion based on your experimental questions.

2

Step 2: Search Available Cre Lines

Use resources like the Jackson Laboratory Cre Portal, MGI, or IMSR to identify candidate lines expressing in your tissue of interest.

3

Step 3: Review Published Characterization

For each candidate, review published data on expression pattern, recombination efficiency, and any reported issues.

4

Step 4: Evaluate Specificity and Timing

Assess whether expression pattern matches your needs. Consider both intended and unintended expression.

5

Step 5: Check Availability and Background

Verify the line is available, and consider whether strain background is compatible with your floxed allele.

6

Step 6: Plan Validation Experiments

Plan to validate Cre expression and recombination in your specific cross using reporter alleles or PCR based methods.

Common Selection Mistakes

Mistake: Relying solely on promoter name

Solution: Always check actual expression data. Transgenic Cre lines often have expression patterns that differ from the endogenous gene due to position effects.

Mistake: Ignoring germline recombination

Solution: Some Cre lines show germline activity, especially in females. Always breed Cre through males for the first generation and genotype for recombination.

Mistake: Not considering developmental expression

Solution: A Cre that is specific in adults may have broader expression during development. If your gene is essential during development, this can cause unexpected lethality.

Mistake: Overlooking Cre toxicity

Solution: High Cre expression can be toxic independent of target gene deletion. Always include Cre only controls in your experiments.

Mistake: Assuming complete deletion

Solution: Most Cre lines do not achieve 100% recombination. Plan for mosaic deletion and consider using reporter alleles to assess efficiency.

Resources for Finding Cre Lines

Jackson Laboratory Cre Portal

Comprehensive database of Cre driver lines with expression data and availability information.

Mouse Genome Informatics (MGI)

Searchable database of recombinase alleles with links to expression data and publications.

International Mouse Strain Resource (IMSR)

Global database of mouse strains including Cre lines, with source information.

Published Literature

Original characterization papers often contain the most detailed expression and recombination data.

What Researchers Say

iTL produced four conditional knockout mouse models on our behalf. They have been extremely helpful and informative at all stages of the project; all the way from construct design to breeding strategies and genotyping the new mouse models. I know where to turn when the needs comes up again for another mouse project; it is certainly faster and cheaper than doing this by ourselves.

William A. Coetzee, DSc

NYU School of Medicine

The Hephaestin flox model ingenious has made for us has been great. It has helped generate eight research publications.

Joshua Dunaief, PhD, MD

University of Pennsylvania

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Need Help Selecting a Cre Line?

Our scientific consultants can help you identify the optimal Cre driver for your conditional knockout project. We can also assist with breeding strategies and validation approaches.

Frequently Asked Questions

Start by defining your tissue, cell type, and timing requirements. Search databases like Jackson Laboratory Cre Portal or MGI for candidate lines. Review published characterization data for expression pattern, recombination efficiency, and any reported issues. Verify availability and strain background compatibility. Plan validation experiments using reporter alleles.

Constitutive Cre lines express continuously based on their promoter, causing recombination as soon as the promoter is active (often during development). Inducible Cre lines (CreERT2, Tet-inducible) require an external inducer (tamoxifen, doxycycline) to activate recombination, allowing temporal control over when deletion occurs.

Some Cre lines show activity in the germline, which can cause recombination in all offspring tissues regardless of the Cre's intended tissue specificity. This effectively converts your conditional allele to a conventional knockout. To avoid this, breed Cre through males for the first generation and genotype for unexpected recombination.